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They're human... they couldn't really be anything else.

1) all of the traits they share with humans: able to use the same weapons, wear the same kind of uniforms, etc.
2) in the e3 binks from 2003 Barney Calhoun is wearing a combine uniform [without the mask]
3) they bleed red

That's just my theory, and if this has been mentioned before.. well...
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